And the protection of the clan/group, yes true.
It's overall aggression for a variety of purposes in my view.
The problem is, the testosterone is still there but the reasons for it existing have largely been made obselete, other than a few select uses.
Testosterone drives the sexual urge which as I think you said earlier, is an ever present hunger in men, generally speaking.
Women, especially in western society are no longer dependent in the same ways they were. Men don't have the same stranglehold on providing or protecting.
I get my own food and provide my own shelter. For me and my offspring. This is very threatening to men. It means I can deny them sex.
If I need protection, I call the police. Or get a dog (simplistically). The need for me to have a man is zero.
Men don't like that.
There are also very useful studies on men finding successful women threatening to their self esteem.
m.huffingtonpost.co.uk/entry/successful-girlfriend-men-threatened_n_3838295
This is where alot of the anger comes from in my opinion. The incel stuff is very telling which I'm sure you know about.
They have the aggressive urges but no longer the entitlement to act on them as they would have had or as chimps have. Doesnt stop them trying or being violent though. Add in the weird sexual and emotional dysfunction stuff I mentioned earlier and it's a toxic cocktail.