That article obscures the issue. It's not clear at all. It goes into great detail about DSDs without emphasising the fact that most people want to know - the two boxers in question are male.
That you've come away less clear on this fact than before you read the article is exactly their intent.
The stuff about the SRY gene is interesting but irrelevant - the boxers don't have Swyer syndrome. We can plainly see that as it's obvious both have gone through male puberty, plus this can be surmised from what has been said (and not said) by the IOC and the IBA.
The article selectively quotes Dr Emma Hilton, misrepresenting her position. It makes it sound like she wants more testing as it's ambiguous. But in fact, she is very clear that she thinks the boxers are male.
Here is a thread that she has retweeted, saying "The only thread you’ll ever need on the IOC, DSDs and male advantage."
https://x.com/TLexercise/status/1819653748840448310
I strongly suggest everyone reads it, it's much clearer than that article.
The boxers likely have the male DSD 5ARD, same as Caster Semanya.
Which means that although their genitals will have been underdeveloped at birth, leading to them being incorrectly identified as female, male puberty will have hit at the normal.time, at which point their penis will have become obvious, alongside all the other changes typical.of male puberty.
Meaning they do have the SRY gene, so the discussion about that is irrelevant and is creating confusion.
They will have known they are make since puberty.
The quote in the BBC article above a 33 year old discovering she is XY is totally irrelevant to this discussion as she will have had a different DSD, likely Swyer syndrome. Again, confusing the issue.
The BBC is under immense pressure from the trams rights lobby to not speak plainly on this. It's a shame, but their reporting really can't be trusted on sex and gender related issues. (Even when it's actually nothing to do with trans people).