Well, I've got an older model Bee and a Cameleon and my vote goes to the Cameleon!
I think both of them are fantastic (hence owning both!) - fantastic designs, both steer beautifully etc but for long walks, especially over rougher terrain, the Cam wins hands down. It is bigger and bulkier than the Bee, but as you say you've got plenty of room in the car, it really shouldn't be an issue for you.
The Bee is a fair bit lighter and is a bit easier to steer one handed - when out with small baby whilst holding a lively toddlers hand for example! - so great for shopping etc, but if on an uneven surface, I definitely notice the difference in how smoothly it pushes. Folds lovely and small though compared to the Cam.
The carrycot bit on the Cam is nice and spacious - used for both DS and DD - long AND heavy babies!- until about four months old. The only reason we changed to the pram seat early is because both children hated not being able to see out, once we changed over, trips out were much less stressful! We got around the problem of the bucket seat not being quite flat enough for a small baby by using those foam wedge things that are used in a car seat for newborns, it changed the bucket style seat into a nice gentle curve, much easier on babys back!
So, there you have it...Cameleon wins my vote, had it for 3 1/2 years and it is still going strong. We've also had fantastic customer service from Bugaboo in the past.
HTH and sorry for the ramble ! 