To me, it means misbehaving, of a child or a machine. eg "Dd1 has been acting up all morning" or "My washing machine's been acting up"
At the college where I work, everyone persistently uses it to mean "acting", as in taking up a post temporarily when someone is on maternity or sick leave, or a post hasn't been filled yet. So they say
"An acting up arrangement has arisen for a blah blah blah. The acting up will be for one term, until blah blah."
Has anyone else heard of this? Please agree with me and say it's just ugly and wrong. It's "Acting Head of blah blah blah"....isn't it?